William R. Kenan Professor of Biology Emeritus


(ORCID number = 000-0002-9438-103X)


The Nature HandbookWilliams, E.H., Jr. (ed.) 2012. Pathways to Excellence in Teaching. Couper Press, Clinton. NY. 81 pp. (softcover) ISBN 978-1-937370-06-0

Williams, E.H., Jr. 2005. The Nature Handbook: A Guide to Observing the Great Outdoors. Oxford Univ. Press, NY. 263 pp. (softcover) ISBN 0-19-517194-2 

Stokes, D., L. Stokes, & E. Williams. 1991. Stokes Butterfly Book. Little, Brown & Co., Boston. 96 pp. ISBN 0-316-81780-5

Williams, E.H. (ed.). 1990. A Marsh for All Seasons: Information and Teacher's Guide for Utica Marsh. Utica Marsh Council, Utica, NY

Selected Articles and Chapters

(click on pdf to download a copy)

Williams, E.H. 2019. The butterflies (Lepidoptera) of an isolated island: Monhegan, Maine. Northeastern Naturalist 26(3):537-544.pdf

Oberhauser, K.S., A. Alonso, S.B. Malcolm, E.H. Williams, and M.P. Zalucki. 2019. Lincoln Brower, champion for monarchs. Frontiers in Ecology and Evolution 8 May 2019. pdf

Brower, L.P., E.H. Williams, et al. 2018. A long-term survey of spring monarch butterflies in north-central Florida. Journal of Natural History 52:2025-2046. Please email me for a copy.

Brower, L.P., E.H. Williams, P Jaramillo-Lopez, D.R. Kust, D.A. Slayback, & M.I. Ramirez. 2017. Butterfly mortality and salvage logging from the March 2016 storm in the Monarch Butterfly Biosphere Reserve in Mexico. American Entomologist 63(3):151-164. pdf

Zipf, L., E.H. Williams, R.B. Primack, & S. Stichter. 2017. Climate effects on late-season flight times of Massachusetts butterflies. International Journal of Biometeorology DOI 10.1007/s00484-017-1347-8. Email me for a copy or available at https://link.springer.com/article/10.1007%2Fs00484-017-1347-8

Alcock, J., L.P. Brower, & E.H. Williams. 2016. Monarch butterflies use regenerating milkweeds for reproduction in mowed hayfields in northern Virginia. Journal of the Lepidopterists' Society 70:177-181. pdf

Pleasants, J.M., E.H. Williams, L.P. Brower, K.S. Oberhauser, & O.R. Taylor. 2016. Conclusion of no decline in summer monarch population not supported. Annals of the Entomological Society of America, online doi: 10.1093/aesa/sav115. pdf

Brower, L.P., D.A. Slayback, P. Jaramillo-Lopez, I. Ramirez, K.S. Oberhauser, E.H. Williams, & L.S. Fink. 2016. Illegal logging of ten hectares of forest in the Sierra Chincua monarch butterfly overwintering arera in Mexico. American Entomologist (summer, 2016):92-97. pdf

Williams, E.H., & L.P. Brower. 2016. A conservation concern: how many monarchs are there? News of the Lepidopterists' Society 58:90-93. pdf

Williams, E.H., & L.P. Brower. 2015. Microclimatic protection of overwintering monarch butterflies provided by Mexico's high elevation Oyamel fir forests: a review. Chap. 9 In: K. Oberhauser,S. Altizer, & K. Nail, eds., Monarchs in a Changing World: Biology and Conservation of an Iconic Insect, Cornell Univ. Press, Ithaca, NY. ISBN 978-0-8014-5315-1. pdf

Fischer, S.J., E.H. Williams, L.P. Brower, & P.A. Palmiotto. 2015. Enhancing monarch butterfly reproduction by mowing fields of common milkweed. American Midland Naturalist 173(2):229-240. pdf

Williams, E.H., S.B. Stichter, C. Hitchcock, C.A. Polgar, & R.B. Primack. 2014. Phenological advancement of Lycaenid butterflies in Massachusetts. Journal of the Lepidopterists' Society 68:167-174. pdf

Polgar, C.A., R.B. Primack, E.H. Williams, S. Stichter, & C. Hitchcock. 2013. Climate effects on the flight period of Lycaenid butterflies in Massachusetts. Biological Conservation 160:25-31. Please email me for a copy.

Williams, E.H. 2012. Population loss and gain in the rare butterfly Euphydryas gillettii (Nymphalidae). Journal of the Lepidopterists' Society 66:147-155. pdf

Williams, E.H. 2012. Epilogue: what students tell us. pp. 77-81 in E.H. Williams (ed.), Pathways to Excellence in Teaching, Couper Press, Clinton, NY. pdf

Van Hook, T., E.H. Williams, L.P. Brower, S. Borkin, & J. Hein. 2012. A standardized protocol for ruler-based measurement of wing length in monarch butterflies, Danaus plexippus L. (Nymphalidae, Danainae). Tropical Lepidoptera Research 22(1):42-52. pdf

Brower, L.P., O.R. Taylor, & E.H. Williams. 2012. Response to Davis: choosing relevant evidence to assess monarch population trends. Insect Conservation and Diversity 5:327-329. (published on-line October, 2011; doi: 10.1111/j.1752-4598.2011.00176.x). pdf [The definitive version is available at www.blackwell-synergy.com.]

Williams, E.H. 2012. Long term effects of climate on two pond predators. American Midland Naturalist 167:336-343. pdf

Brower, L.P., O.R. Taylor, E.H. Williams, D.A. Slayback, R.R. Zubieta, & M.I. Ramirez. 2012. Decline of monarch butterflies overwintering in Mexico: Is the migratory phenomenon at risk? Insect Conservation and Diversity 5(2):95-100. (published on-line March, 2011; doi: 10.1111/j.1752-4598.2011.00142.x). pdf [The definitive version is available at www.blackwell-synergy.com.]

Brower, L.P., E.H. Williams, L.S. Fink, D.A. Slayback, M.I. Ramirez, M.I. Limon Garcia, R.R. Zubieta, S.B. Weiss, W.H. Calvert, & W. Zuchowski. 2011. Overwintering clusters of the monarch butterfly coincide with the least hazardous vertical temperatures in the oyamel forest. Journal of the Lepidopterists' Society 65(1):27-46. pdf

Williams, E.H., P.D. Reynolds, & O. Oerlemans. 2010. Interdisciplinary teaching about the Adirondacks. In: P. Cenkl (ed.), Nature and Culture in the Northern Forest, Univ. Iowa Press. pdf

Brower, L.P., E.H. Williams, D.A. Slayback, L.S. Fink, M.I. Ramirez, R.R. Zubieta, M.I. Limon Garcia, P. Gier, J.A. Lear, & T. van Hook. 2009. Oyamel fir forest trunks provide thermal advantages for overwintering monarch butterflies in Mexico. Insect Conservation and Diversity 2:163-175. pdf

Pfitsch, W.A., & E.H. Williams. 2009. Habitat restoration for lupine and specialist butterflies. Restoration Ecology 17(2):226-233. pdf

Brower, L.P., E.H. Williams, L.S. Fink, R.R. Zubieta Hernandez, & M.I. Ramirez. 2008. Monarch butterfly clusters provide microclimatic advantages during the overwintering season in Mexico. Journal of the Lepidopterists' Society 62:177-188. pdf

Boggs, C.L., C.E. Holdren, I.G. Kulahci, T.C. Bonebrake, B.D. Inouye, J.P. Fay, A. MacMillan, E.H. Williams, & P.R. Ehrlich. 2006. Delayed population explosion of an introduced butterfly. Journal of Animal Ecology 75:466-475. pdf

Kocher, S.D., & E.H. Williams. 2000. The diversity and abundance of North American butterflies vary with habitat disturbance and geography. Journal of Biogeography 27(4):785-794. pdf

Knight, A.L., Brower, L.P., & E.H. Williams. 1999. Spring remigration of the monarch butterfly, Danaus plexippus (Lepidoptera: Nymphalidae) in north-central Florida: estimating population parameters using mark-recapture. Biological Journal of the Linnaean Society 68:531-556. pdf

Brooks, J.S., E.H. Williams, & P. Feeny. 1996. Quantification of contact oviposition stimulants for Black Swallowtail butterfly, Papilio polyxenes, on the leaf surfaces of wild carrot, Daucus carota. Journal of Chemical Ecology 22(12):2341-2357. pdf

Williams, E.H. 1995. Fire-burned habitat and reintroductions of the butterfly Euphydryas gillettii (Nymphalidae). Journal of the Lepidopterists' Society 49:183-191. pdf

Bowers, M.D., & E.H. Williams. 1995. Variable chemical defence in the checkerspot butterfly Euphydryas gillettii (Lepid.: Nymphalidae). Ecological Entomology 20:208-212. pdf

Bart, K.M., & E.H. Williams. 1993. Use of dental wax for the study of insect behavior by scanning electron microscopy. Microscopy Research and Technique 26:180-181. pdf

Williams, E.H. 1990. Dietary breadth in Euphydryas gillettii (Nymphalidae). Journal of the Lepidopterists' Society 44(2):94-95. pdf

Williams, E.H. 1988. Habitat and range of Euphydryas gillettii (Nymphalidae). Journal of the Lepidopterists' Society 42(1):37-45. pdf

Williams, E.H., & M.D. Bowers 1987. Factors affecting host-plant use by the montane butterfly Euphydryas gillettii (Nymphalidae). American Midland Naturalist 118(1):153-161. pdf

McQuade, D.B., E.H. Williams, & H.B. Eichenbaum. 1986. Cues used for localizing food by the Gray Squirrel (Sciurus carolinensis). Ethology 72(1):22-30. pdf

Williams, E.H., C.E. Holdren, & P.R. Ehrlich. 1984. The life history and ecology of Euphydryas gillettii Barnes (Nymphalidae). Journal of the Lepidopterists' Society 38(1):1-12. pdf

Williams, E.H., E.L. Mather, & S.M. Carter. 1984. Toxicity of the herbicides endothall and diquat to benthic crustacea. Bulletin of Environmental Contamination and Toxicology 33(4):418-422. pdf

Williams, E.H. 1981. Thermal influences on oviposition in the montane butterfly Euphydryas gillettii. Oecologia 50(3):342-346. pdf

Williams, E.H. 1980. Disjunct distributions of two aquatic predators. Limnology and Oceanography 25(6):999-1006. pdf


Selected Non-Technical Publications

Williams, E.H. 2012. Microclimates and the survival of overwintering monarchs. American Butterflies 20(2):8-13. pdf

Williams, E.H. 2009. Lifestyles of the scaled and beautiful: Pearl and Northern Crescents. American Butterflies 17(2):4-13. pdf

Williams, E.H. 2006. Baltimore Checkerspots: Butterflies of privilege. American Butterflies 14(2):4-13. pdf

Williams, E.H. 2002. Harris' Checkerspot: A very particular butterfly. American Butterflies 10(2):18-25. pdf

Willaims, E.H. 2001. Gillett's Checkerspot: Life in a mountain meadow. American Butterflies 9(2):6-13 pdf

Back to Top